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Reach

a fatty acyl-CoA

362,296 routes · scope human · up to 3 steps · built 5 October 2026 · Reactions · API

Inferred

Every route here is a hypothesis drawn from reference reactions (Rhea and Human-GEM), not a finding. Currency species such as water, ATP and NAD(H) never form a step. Evidence key

339 compounds within 3 steps: 10 in 1, 44 in 2, 285 in 3

Does it reach a compound?

Two steps

an N-acyl-(4R)-4-hydroxysphinganine-1-phosphoethanolamineReach from here →
1. a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
2. an N-acyl-(4R)-4-hydroxysphinganine + a 1,2-diacyl-sn-glycero-3-phosphoethanolamine = an N-acyl-(4R)-4-hydroxysphinganine-1-phosphoethanolamine + a 1,2-diacyl-sn-glycerol
Catalysed by LPGAT1, SGMS2, SAMD8
1,2-dihexadecanoyl-sn-glycerolReach from here →
1. sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
2. 1,2-dihexadecanoyl-sn-glycero-3-phosphocholine + an N-acylsphing-4-enine = 1,2-dihexadecanoyl-sn-glycerol + a sphingomyelin
Catalysed by CERS1, CERS6, CERS3, CERS5 and 3 more
1-(9Z-octadecenoyl)-2-acyl-sn-3-glycerolReach from here →
1. sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
2. 1-(9Z-octadecenoyl)-2-acyl-sn-3-glycerol + a sphingomyelin = a 1-(9Z-octadecenoyl)-2-acyl-sn-glycero-3-phosphocholine + an N-acylsphing-4-enine
Catalysed by CERS1, CERS6, CERS3, CERS5 and 4 more
1-O-acyl-N-(acetyl)-sphing-4-enineReach from here →
1. a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
2. N-(acetyl)-sphing-4-enine + a 1,2-diacyl-sn-glycero-3-phosphoethanolamine = 1-O-acyl-N-(acetyl)-sphing-4-enine + a 1-acyl-sn-glycero-3-phosphoethanolamine
Catalysed by LPGAT1, PLA2G15One of 2 routes

Three steps

CHEBI:1146900Reach from here →
Catalysed by CERS1, CERS6, CERS3, CERS5 and 10 moreGolgi apparatus, mitochondrial matrixOne of 46 routes
N(omega)-methyl-L-arginineReach from here →

Page 2 of 7

Catalysed by GLYATL2, AGXT2

AReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
a beta-D-glucosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-glucoside + an N-acylsphing-4-enine
cholesterol + AH2 + O2 = 25-hydroxycholesterol + A + H2O

Catalysed by CERS1, CERS6, CERS3, CERS5 and 5 more

an acyl-4'-phosphopantetheineReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
an acyl-CoA + H2O = an acyl-4'-phosphopantetheine + adenosine 3',5'-bisphosphate + 2 H(+)

Catalysed by CERS1, CERS6, CERS3, CERS5 and 9 moreGolgi apparatus

glycoursodeoxycholateReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
glycine to glycoursodeoxycholate conversion

Catalysed by GLYATL2cytosolOne of 2 routes

N-(9Z-octadecenoyl)glycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
(9Z)-octadecenoyl-CoA + glycine = N-(9Z-octadecenoyl)glycine + CoA + H(+)

Catalysed by GLYATL2, GLYATL3

N-octadecanoylglycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
octadecanoyl-CoA + glycine = N-octadecanoylglycine + CoA + H(+)

Catalysed by GLYATL2

N-(9Z-hexadecenoyl)-glycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
(9Z)-hexadecenoyl-CoA + glycine = N-(9Z-hexadecenoyl)-glycine + CoA + H(+)

Catalysed by GLYATL2

cholesterol sulfateReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
a beta-D-glucosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-glucoside + an N-acylsphing-4-enine
cholesterol + 3'-phosphoadenylyl sulfate = cholesterol sulfate + adenosine 3',5'-bisphosphate + H(+)

Catalysed by CERS1, CERS6, CERS3, CERS5 and 6 more

(11Z,14Z,17Z-eicosatrienoyl)-cholesterolReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
a beta-D-glucosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-glucoside + an N-acylsphing-4-enine
1-hexadecanoyl-2-(11Z,14Z,17Z-eicosatrienoyl)-sn-glycero-3-phosphocholine + cholesterol = (11Z,14Z,17Z-eicosatrienoyl)-cholesterol + 1-hexadecanoyl-sn-glycero-3-phosphocholine

Catalysed by CERS1, CERS6, CERS3, CERS5 and 5 more

a 3-hydroxyacyl derivative of bacterial toxinReach from here →

a fatty acyl-CoA + H2O = a fatty acid + CoA + H(+)
a 3-(acyloxy)acyl derivative of bacterial toxin + H2O = a 3-hydroxyacyl derivative of bacterial toxin + a fatty acid + H(+)
a 3-(acyloxy)acyl derivative of bacterial toxin + H2O = a 3-hydroxyacyl derivative of bacterial toxin + a fatty acid + H(+)

Catalysed by ACOT13, AOAH

N-hexadecanoylglycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
hexadecanoyl-CoA + glycine = N-hexadecanoylglycine + CoA + H(+)

Catalysed by GLYATL2, GLYATL3

N-dodecanoylglycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
dodecanoyl-CoA + glycine = N-dodecanoylglycine + CoA + H(+)

Catalysed by GLYATL2

N-decanoylglycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
decanoyl-CoA + glycine = N-decanoylglycine + CoA + H(+)

Catalysed by GLYATL2

N-octanoylglycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
octanoyl-CoA + glycine = N-octanoylglycine + CoA + H(+)

Catalysed by GLYATL2

N(1)-(5-phospho-beta-D-ribosyl)glycinamideReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
5-phospho-beta-D-ribosylamine + glycine + ATP = N(1)-(5-phospho-beta-D-ribosyl)glycinamide + ADP + phosphate + H(+)

Catalysed by GLYATL2, GART

L-cystine-bis-glycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
L-cystine-bis-glycine + 2 H2O = L-cystine + 2 glycine

Catalysed by GLYATL2, DPEP1, DPEP2

sn-glycero-3-phosphoethanolamineReach from here →

a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
N-(acetyl)-sphing-4-enine + a 1,2-diacyl-sn-glycero-3-phosphoethanolamine = 1-O-acyl-N-(acetyl)-sphing-4-enine + a 1-acyl-sn-glycero-3-phosphoethanolamine
a 1-acyl-sn-glycero-3-phosphoethanolamine + H2O = sn-glycero-3-phosphoethanolamine + a fatty acid + H(+)

Catalysed by LPGAT1, PLA2G15, ASPG

a 2-acyl-6-[alpha-D-mannosyl-(1->2)-alpha-D-mannosyl-(1->6)-2-phosphoethanolamine-alpha-D-mannosyl-(1->4)-alpha-D-glucosaminyl]-1-(1-radyl,2-acyl-sn-glycero-3-phospho)-1D-myo-inositolReach from here →

a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
a 2-acyl-6-[alpha-D-mannosyl-(1->6)-alpha-D-mannosyl-(1->4)-alpha-D-glucosaminyl]-1-(1-radyl,2-acyl-sn-glycero-3-phospho)-1D-myo-inositol + a 1,2-diacyl-sn-glycero-3-phosphoethanolamine = a 2-acyl-6-[alpha-D-mannosyl-(1->6)-2-phosphoethanolamine-alpha-D-mannosyl-(1->4)-alpha-D-glucosaminyl]-1-(1-radyl,2-acyl-sn-glycero-3-phospho)-1D-myo-inositol + a 1,2-diacyl-sn-glycerol
a 2-acyl-6-[alpha-D-mannosyl-(1->6)-2-phosphoethanolamine-alpha-D-mannosyl-(1->4)-alpha-D-glucosaminyl]-1-(1-radyl,2-acyl-sn-glycero-3-phospho)-1D-myo-inositol + a di-trans,poly-cis-dolichyl beta-D-mannosyl phosphate = a 2-acyl-6-[alpha-D-mannosyl-(1->2)-alpha-D-mannosyl-(1->6)-2-phosphoethanolamine-alpha-D-mannosyl-(1->4)-alpha-D-glucosaminyl]-1-(1-radyl,2-acyl-sn-glycero-3-phospho)-1D-myo-inositol + a di-trans,poly-cis-dolichyl phosphate + H(+)

Catalysed by LPGAT1, PIGN, PIGB

glycyldehydrophenylalanineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
glycyldehydrophenylalanine + H2O = 2,3-didehydrophenylalanine + glycine

Catalysed by GLYATL2, DPEP1

N-(9Z,12Z-octadecadienoyl)-glycineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
(9Z,12Z)-octadecadienoyl-CoA + glycine = N-(9Z,12Z-octadecadienoyl)-glycine + CoA + H(+)

Catalysed by GLYATL2

cholesteryl (7Z)-octadecenoateReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
a beta-D-glucosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-glucoside + an N-acylsphing-4-enine
(7Z)-octadecenoyl-CoA + cholesterol = cholesteryl (7Z)-octadecenoate + CoA

Catalysed by CERS1, CERS6, CERS3, CERS5 and 5 more

(7Z)-octadecenoyl-CoAReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
acyl-CoA pool formation and breakdown

Catalysed by CERS1, CERS6, CERS3, CERS5 and 6 moreGolgi apparatus, cytosol

acetaldehydeReach from here →

a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + L-serine = a 1,2-diacyl-sn-glycero-3-phospho-L-serine + ethanolamine
ethanolamine ammonia-lyase (acetaldehyde-forming)

Catalysed by LPGAT1, PTDSS1, PTDSS2cytosolOne of 2 routes

cholineReach from here →

a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + a 1,2-diacyl-sn-glycero-3-phosphocholine = an N-acyl-1,2-diacyl-sn-glycero-3-phosphoethanolamine + a 2-acyl-sn-glycero-3-phosphocholine + H(+)
a 2-acyl-sn-glycero-3-phosphocholine + H2O = a 2-acyl-sn-glycerol 3-phosphate + choline + H(+)

Catalysed by LPGAT1, PLA2G4E, PLAAT5, PLAAT2 and 1 moreOne of 2 routes

arachidonoyl-CoAReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
acyl-CoA pool formation and breakdown

Catalysed by CERS1, CERS6, CERS3, CERS5 and 6 moreGolgi apparatus, cytosol

palmitoyl-CoAReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
acyl-CoA pool formation and breakdown

Catalysed by CERS1, CERS6, CERS3, CERS5 and 6 moreGolgi apparatus, cytosol

oleoyl-CoAReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
acyl-CoA pool formation and breakdown

Catalysed by CERS1, CERS6, CERS3, CERS5 and 6 moreGolgi apparatus, cytosol

CHEBI:15570Reach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of D-alanine and glycine (cytosol to extracellular)

Catalysed by GLYATL2, SLC3A2, SLC7A10extracellular space, cytosolOne of 2 routes

L-leucineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of glycine and leucine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A5extracellular space, cytosolOne of 5 routes

CHEBI:15611Reach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
S-Adenosyl-L-methionine:glycine N-methyltransferase

Catalysed by GLYATL2, GNMTcytosol

CHEBI:15699Reach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of glycine and homoserine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A5extracellular space, cytosolOne of 4 routes

hexadecanoic acidReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
N-Acylsphingosine amidohydrolase
(10Z)-heptadecenoic acid formation

Catalysed by CERS1, CERS6, CERS3, CERS5 and 3 morelysosome, extracellular spaceOne of 12 routes

1-acylglycerone 3-phosphateReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
Acyl-CoA:glycerone-phosphate O-acyltransferase

Catalysed by CERS1, CERS6, CERS3, CERS5 and 7 moreGolgi apparatus, cytosol

a sterolReach from here →

a 2-acyl-sn-glycero-3-phosphoethanolamine + a fatty acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + CoA
a 1,2-diacyl-sn-glycero-3-phosphoethanolamine + a 1,2-diacyl-sn-glycero-3-phosphocholine = an N-acyl-1,2-diacyl-sn-glycero-3-phosphoethanolamine + a 1-acyl-sn-glycero-3-phosphocholine + H(+)
a sterol + a 1,2-diacyl-sn-glycero-3-phosphocholine = a sterol ester + a 1-acyl-sn-glycero-3-phosphocholine

Catalysed by LPGAT1, PLAAT5, PLAAT2, LCAT

CHEBI:15971Reach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of glycine and histidine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A6extracellular space, cytosolOne of 2 routes

phosphatidylethanolamineReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
Acyl-CoA:1-acyl-sn-glycero-3-phosphoethanolamine O-acyltransferase

Catalysed by CERS1, CERS6, CERS3, CERS5 and 11 moreGolgi apparatus, cytosolOne of 2 routes

1,2-diacyl-sn-glycero-3-phosphocholine(1+)Reach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
Acyl-CoA:1-acyl-sn-glycero-3-phosphocholine O-acyltransferase

Catalysed by CERS1, CERS6, CERS3, CERS5 and 10 moreGolgi apparatus, cytosolOne of 2 routes

11-cis-retinolReach from here →

a fatty acyl-CoA + H2O = a fatty acid + CoA + H(+)
an all-trans-retinyl ester + H2O = 11-cis-retinol + a fatty acid + H(+)
all-trans-retinyl ester acylhydrolase, 11-cis-retinol forming

Catalysed by ACOT13, RPE65cytosolOne of 2 routes

phosphatidic acidReach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
acyl-CoA:sphingosine N-acyltransferase
acyl-CoA:1-acyl-sn-glycerol-3-phosphate 2-O-acyltransferase

Catalysed by CERS1, CERS6, CERS3, CERS5 and 15 moreGolgi apparatus, mitochondrial matrixOne of 9 routes

L-citrullineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of citrulline and glycine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A9, SLC3A1, SLC7A6extracellular space, cytosol

L-valineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of glycine and valine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A5extracellular space, cytosolOne of 4 routes

CHEBI:16436Reach from here →

sphing-4-enine + a fatty acyl-CoA = an N-acylsphing-4-enine + CoA + H(+)
ceramide:phosphatidylcholine cholinephosphotransferase
ceramide cholinephosphotransferase (CDP-choline)

Catalysed by CERS1, CERS6, CERS3, CERS5 and 4 moreGolgi apparatus, endoplasmic reticulum

L-arginineReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of arginine and glycine (cytosol to extracellular)

Catalysed by GLYATL2, SLC7A6extracellular space, cytosolOne of 2 routes

CHEBI:16523Reach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
transport of D-serine and glycine (cytosol to extracellular)

Catalysed by GLYATL2, SLC3A2, SLC7A10extracellular space, cytosolOne of 2 routes

protein polypeptide chainReach from here →

a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
a fatty acyl-CoA + glycine = an N-(fatty acyl)-glycine + CoA + H(+)
PROTEIN_BS

Catalysed by GLYATL2cytosol