a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine

362,296 routes · scope human · up to 3 steps · built 5 October 2026 · Reactions · API

185 compounds within 3 steps: 2 in 1, 16 in 2, 167 in 3

Three steps

dodecanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
cholesteryl 3-beta-D-galactosideReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. a beta-D-galactosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-galactoside + an N-acylsphing-4-enine
Catalysed by GBA1, GBA3, GBA2One of 3 routes
(11Z)-docosenoyl-CoA(4-)Reach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
(7Z)-tetradecenoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
docosanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
octadecanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
alpha-linolenic acidReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
elaidic acidReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosolOne of 13 routes
alpha-D-galactoseReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. UDP-alpha-D-galactose:N-acylsphingosine D-galactosyltransferase
3. D-galactosyl-N-acylsphingosine galactohydrolase
Catalysed by GBA1, GBA3, UGT8, GALCcytosol, lysosome
all-cis-docosa-4,7,10,13,16,19-hexaenoic acidReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
phosphocholineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. ceramide:phosphatidylcholine cholinephosphotransferase
3. a sphingomyelin + H2O = phosphocholine + an N-acylsphing-4-enine + H(+)
Catalysed by GBA1, GBA3, SGMS1, SGMS2 and 5 moreGolgi apparatus
tetradecanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(9Z)-octadecenoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
hexacosanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
tetracosanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
an N-acylsphinganineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. sphingosine N-acyltransferase (sphinganine)
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 8 moreGolgi apparatus, cytosolOne of 3 routes
an N-acyl-(4R)-4-hydroxysphinganineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. ceramide:phosphatidylcholine cholinephosphotransferase
3. an N-acyl-(4R)-4-hydroxysphinganine + a 1,2-diacyl-sn-glycero-3-phosphocholine = an N-acyl-(4R)-4-hydroxysphinganine-phosphocholine + a 1,2-diacyl-sn-glycerol
Catalysed by GBA1, GBA3, SGMS1, SGMS2Golgi apparatus
eicosanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(9Z)-hexadecenoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(15Z)-tetracosenoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(5Z,8Z,11Z,14Z)-eicosatetraenoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(11Z)-eicosenoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. N-Acylsphingosine amidohydrolase
3. (10Z)-heptadecenoic acid formation
Catalysed by GBA1, GBA3, ASAH1lysosome, extracellular spaceOne of 12 routes
(24S)-hydroxycholesterolReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. cholesterol + reduced [NADPH--hemoprotein reductase] + O2 = (24S)-hydroxycholesterol + oxidized [NADPH--hemoprotein reductase] + H2O + H(+)
Catalysed by GBA1, GBA3, CYP46A1
a fatty aldehydeReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. an N-acylsphing-4-enine + H2O = sphing-4-enine + a fatty acid
3. a fatty aldehyde + NAD(+) + H2O = a fatty acid + NADH + 2 H(+)
Catalysed by GBA1, GBA3, NAAA, ASAH1 and 5 more
cholesteryl hexadecanoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. cholesterol + hexadecanoyl-CoA = cholesteryl hexadecanoate + CoA
Catalysed by GBA1, GBA3, SOAT1
a 1-O-alkyl-2-acyl-sn-glycero-3-phosphocholineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. Acyl-CoA:1-alkyl-sn-glycero-3-phosphocholine O-acyltransferase
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 6 moreGolgi apparatus, cytosol
sphingosine 1-phosphateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. ATP:sphingosine 1-phosphotransferase
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 5 moreGolgi apparatus, cytosolOne of 2 routes
D-galactoseReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. an N-acylsphing-4-enine + UDP-alpha-D-galactose = a beta-D-galactosyl-(1<->1')-N-acylsphing-4-enine + UDP + H(+)
3. a beta-D-galactosyl-(1<->1')-N-acylsphing-4-enine + H2O = D-galactose + an N-acylsphing-4-enine
Catalysed by GBA1, GBA3, UGT8, GALC and 1 more
cholesteryl (9Z,12Z)-octadecadienoateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. (9Z,12Z)-octadecadienoyl-CoA + cholesterol = cholesteryl (9Z,12Z)-octadecadienoate + CoA
Catalysed by GBA1, GBA3, SOAT1One of 2 routes
D-glucoseReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. UDP-glucose:N-acylsphingosine D-glucosyltransferase
3. D-Glucosyl-N-acylsphingosine glucohydrolase
Catalysed by GBA1, GBA3, UGCG, GBA2cytosolOne of 5 routes
25-hydroxycholesterolReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. cholesterol + AH2 + O2 = 25-hydroxycholesterol + A + H2O
Catalysed by GBA1, GBA3, CH25HOne of 3 routes
N-octanoylsphing-4-enineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. beta-D-glucosyl-N-octanoylsphing-4E-enine + cholesterol = N-octanoylsphing-4-enine + cholesteryl 3-beta-D-glucoside
Catalysed by GBA1, GBA3
cholesteryl (9Z-octadecenoate)Reach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. cholesterol + (9Z)-octadecenoyl-CoA = cholesteryl (9Z-octadecenoate) + CoA
Catalysed by GBA1, GBA3, SOAT2, SOAT1One of 2 routes
N-(acetyl)-sphing-4-enineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. 1-O-hexadecyl-2-acetyl-sn-glycero-3-phosphocholine + sphing-4-enine = 1-O-hexadecyl-sn-glycero-3-phosphocholine + N-(acetyl)-sphing-4-enine + H(+)
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 3 moreGolgi apparatusOne of 2 routes
a 1,3-diacylglycerolReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. a 1-acylglycerol + an acyl-CoA = a 1,3-diacylglycerol + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 4 moreGolgi apparatus
a 1,2-diacylglycerolReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. a 2-acylglycerol + an acyl-CoA = a 1,2-diacylglycerol + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 7 moreGolgi apparatusOne of 2 routes
(24R)-hydroxycholesterolReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. cholesterol + reduced [NADPH--hemoprotein reductase] + O2 = (24R)-hydroxycholesterol + oxidized [NADPH--hemoprotein reductase] + H2O + H(+)
Catalysed by GBA1, GBA3, CYP3A4
a 1,2-diacyl-sn-glycero-3-phospho-L-serineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. a 1-acyl-sn-glycero-3-phospho-L-serine + an acyl-CoA = a 1,2-diacyl-sn-glycero-3-phospho-L-serine + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 5 moreGolgi apparatus
(6Z,9Z,12Z)-octadecatrienoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
dodecanoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
eicosanoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
(9Z,12Z)-octadecadienoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
tetradecanoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
octadecanoyl-CoAReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. acyl-CoA pool formation and breakdown
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 2 moreGolgi apparatus, cytosol
a 1-acylglycerone 3-phosphateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. dihydroxyacetone phosphate + an acyl-CoA = a 1-acylglycerone 3-phosphate + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 3 moreGolgi apparatus
a 1,2-diacyl-sn-glycero-3-phosphocholineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. a 2-acyl-sn-glycero-3-phosphocholine + an acyl-CoA = a 1,2-diacyl-sn-glycero-3-phosphocholine + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 3 moreGolgi apparatusOne of 9 routes
an N-acylglycineReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. an acyl-CoA + glycine = an N-acylglycine + CoA + H(+)
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 5 moreGolgi apparatus
a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol)Reach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. a 1-acyl-sn-glycero-3-phospho-(1D-myo-inositol) + an acyl-CoA = a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol) + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 4 moreGolgi apparatus
a 1-acyl-sn-glycero-3-phosphateReach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. acyl-CoA:sphingosine N-acyltransferase
3. sn-glycerol 3-phosphate + an acyl-CoA = a 1-acyl-sn-glycero-3-phosphate + CoA
Catalysed by GBA1, GBA3, ASAH1, ACER2 and 6 moreGolgi apparatus
a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol 4-phosphate)Reach from here →
1. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
2. a beta-D-xylosyl-(1<->1')-N-acylsphing-4-enine + cholesterol = cholesteryl 3-beta-D-xyloside + an N-acylsphing-4-enine
3. a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol 4-phosphate)(out) + cholesterol(in) = a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol 4-phosphate)(in) + cholesterol(out)
Catalysed by GBA1, GBA3, OSBPinner side of the membrane, outer side of the membraneOne of 2 routes